Henry Norberto Valdes Marin, 36, alias Pollo – meaning Chicken – and Mauricio Alberto Gonzalez, 44, aka Ronco – meaning Husky.
"These are the most dangerous Colombian citizens to have been detained in Spain up until now due to their history and the variety of criminal activities that they were involved with," said a statement from Spain's interior ministry.
The pair are suspected of working for Colombian drug cartels as "debt collectors". They are also wanted for drug trafficking and illegal weapons possession.
Spanish police said they had relocated to Spain to continue their criminal activities in the country, which serves as the main entry point for South American cocaine into Europe.
Valdes Marin is believed to have acted as logistics chief of La Negra, a group of hit men working for Colombia's Norte del Valle drug cartel.
La Negra is thought to operate across eight countries and is blamed for the murder of more than 200 people in Colombia alone, including the footballer, Albeiro "El Paloma" Usuriaga, who was gunned down in 2004.
Gonzales Sepulveda is one of the suspected leaders of the AUC, a paramilitary organisation serving Colombia's Envigado drug cartel, whose hallmark was chopping up bodies using a chainsaw. Now confined to a wheelchair after taking three bullets in a shoot-out, he had skin grafted to his finger tips to disguise his identity.
The pair, who both appear on Colombia's list of most wanted criminals, were arrested at different address in the Valencia region on Sunday night by Spanish police acting on information given to them by the Colombian counterparts.
They now face extradition to Colombia.
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